Multimorbidity as a predictor of mortality in companion dogs

The authors thank Dog Aging Project participants, their dogs, and community veterinarians, as well as all Dog Aging Project Team members (2025) (https://dogagingproject.org/team-acknowledgements) for their important contributions. The authors would also like to thank Kelsey Patterson and the other members of the Hoffman Lab for all their assistance.

DAP consortium author list:

Joshua M. Akey1, Rozalyn M. Anderson2,3, Elhanan Borenstein4-6, Marta G. Castelhano7,8, Amanda E. Coleman9, Matthew D. Dunbar10, Virginia R. Fajt11, Erica C Jonlin12,13, Matt Kaeberlein12,14, Elinor K. Karlsson15,16, Kathleen F. Kerr17, Jing Ma18, Evan L. MacLean19, Stephanie McGrath20,21, Natasha J Olby22, Daniel E.L. Promislow23,24, May J Reed25, Audrey Ruple26, Stephen M. Schwartz27,28, Sandi Shrager29, Noah Snyder-Mackler30-32, and M. Katherine Tolbert33.

1. Lewis-Sigler Institute for Integrative Genomics, Princeton University, Princeton, NJ, USA

2. University of Wisconsin Madison

3. GRECC William S Middleton Memorial Veterans Hospital, Madison WI

4. Department of Clinical Microbiology and Immunology, Gray Faculty of Medical and Health Sciences, Tel Aviv University, Tel Aviv, Israel

5. Blavatnik School of Computer Science and AI, Tel Aviv University, Tel Aviv, Israel

6. Santa Fe Institute, Santa Fe, NM, USA

7. Cornell Veterinary Biobank, College of Veterinary Medicine, Cornell University, Ithaca, NY, USA

8. Department of Clinical Sciences, College of Veterinary Medicine, Cornell University, Ithaca, NY, USA

9. Department of Small Animal Medicine and Surgery, College of Veterinary Medicine, University of Georgia, Athens, GA, USA

10. Center for Studies in Demography and Ecology, University of Washington, Seattle, WA, USA

11. Department of Veterinary Physiology and Pharmacology, Texas A&M University School of Veterinary Medicine & Biomedical Sciences, College Station, TX, USA

12. Department of Laboratory Medicine and Pathology, University of Washington School of Medicine, Seattle, WA, USA

13. Institute for Stem Cell and Regenerative Medicine, University of Washington, Seattle, WA, USA

14. Department of Oral Health Sciences, University of Washington, Seattle, WA, USA

15. Bioinformatics and Integrative Biology, University of Massachusetts Chan Medical School, Worcester, MA, USA

16. Broad Institute of MIT and Harvard, Cambridge, MA, USA

17. Department of Biostatistics, University of Washington, Seattle, WA, USA

18. Division of Public Health Sciences, Fred Hutchinson Cancer Research Center, Seattle, WA, USA

19. College of Veterinary Medicine, University of Arizona, Tucson, AZ, USA

20. Department of Clinical Sciences, College of Veterinary Medicine and Biomedical Sciences, Colorado State University, Ft. Collins, CO

21. Brain Research Center at Colorado State University

22. Department of Clinical Sciences, College of Veterinary Medicine, North Carolina State University, Raleigh, NC, USA

23. Jean Mayer USDA Human Nutrition Research Center on Aging at Tufts University, Boston, MA

24. Department of Biology, University of Washington, Seattle, WA, USA

25. Department of Medicine, Division of Gerontology and Geriatric Medicine, University of Washington School of Medicine, Seattle, WA

26. Department of Population Health Sciences, Virginia-Maryland College of Veterinary Medicine, Virginia Tech, Blacksburg, VA, USA

27. Epidemiology Program, Fred Hutchinson Cancer Center, Seattle, WA, USA

28. Department of Epidemiology, University of Washington, Seattle, WA, USA

29. Collaborative Health Studies Coordinating Center, Department of Biostatistics, University of Washington, Seattle, WA, USA

30. School of Life Sciences, Arizona State University, Tempe, AZ, USA

31. Center for Evolution and Medicine, Arizona State University, Tempe, AZ, USA

32. School for Human Evolution and Social Change, Arizona State University, Tempe, AZ, USA

33. Department of Small Animal Clinical Sciences, Texas A&M University School of Veterinary Medicine & Biomedical Sciences, College Station, TX, USA

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