These authors contributed equally: Tetsuya Handa, Akihito Harada, Kazumitsu Maehara.

Cell Biology Center, Institute of Innovative Research, Tokyo Institute of Technology, 4259 Nagatsuta, Midori-ku, Yokohama, Japan

Tetsuya Handa & Hiroshi Kimura

Division of Transcriptomics, Medical Institute of Bioregulation, Kyushu University, 3-1-1 Maidashi, Higashi-ku, Fukuoka, Japan

Akihito Harada, Kazumitsu Maehara & Yasuyuki Ohkawa

Institute for Quantitative Biosciences, The University of Tokyo, 1-1-1 Yayoi, Bunkyo-ku, Tokyo, Japan

Shoko Sato & Hitoshi Kurumizaka

Graduate School of Bioscience and Biotechnology, Tokyo Institute of Technology, 4259 Nagatsuta, Midori-ku, Yokohama, Japan

Masaru Nakao, Naoki Goto & Hiroshi Kimura