These authors contributed equally: Judith Miné-Hattab, Marie Villemeur.

Institut Curie, Université PSL, Sorbonne University, CNRS, Nuclear Dynamics, Paris, France

Siyu Liu, Judith Miné-Hattab, Marie Villemeur, Leonid A. Mirny & Angela Taddei

Institute for Integrative Biology of the Cell (I2BC), University of Paris-Saclay, CEA, CNRS, Gif-sur-Yvette, France

Raphaël Guerois

Department of Physics, Massachusetts Institute of Technology, Cambridge, MA, USA

Henrik Dahl Pinholt & Leonid A. Mirny

Institute for Medical Engineering and Science, Massachusetts Institute of Technology, Cambridge, MA, USA

Leonid A. Mirny

Correspondence to Angela Taddei.