aShenzhen Key Laboratory of Metabolic Health, Center for energy metabolism and reproduction, Shenzhen Institutes of Advanced technology, Chinese Academy of Sciences, Shenzhen, China

bSchool of Biological Sciences, University of Aberdeen, Aberdeen, Scotland, UK

cState key laboratory of molecular developmental biology, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing

dCAS Center of Excellence in animal evolution and genetics, Kunming, China

eDepartment of Evolutionary Anthropology

fDuke Global Health Institute, Duke University, Durham, North Carolina, USA

Correspondence to John R. Speakman, Chinese Academy of Sciences Shenzhen Institutes of Advanced Technology, Shenzhen, China. E-mail: [email protected]