Available online 2 December 2023

Brody and Cravchik, 2000, Pfeiffer et al., 2010, Venken et al., 2009.

The authors declare no conflict of interest.

We thank members of the He laboratory for suggestions and discussion during this work.

We thank Bloomington Stock Center and TsingHua Fly Center for fly stocks. Funding: This work was supported by research grants from the National Natural Science Foundation of China (31871461 and 31671510 to H.H.).

There are more references available in the full text version of this article.

© 2023 Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, and Genetics Society of China. Published by Elsevier Limited and Science Press.